thumb|Lullula arborea
The woodlark or wood lark (Lullula arborea) is the only extant species in the lark genus Lullula. It is found across most of Europe, the Middle East, western Asia and the mountains of north Africa. It is mainly resident (non-migratory) in the west of its range, but eastern populations of this passerine bird are more migratory, moving further south in winter.
There are two subspecies of woodlark, L. a. arborea and L. a. pallida. The former is native to northern regions of Europe, while the latter can be found in the south of the woodlark's range. Their diet is mostly composed of seeds but also includes insects during the breeding period. A comparatively small bird, the woodlark is between long and roughly 20% shorter than the skylark. It is a brown bird with a pale underside and has a white-tipped tail.
Taxonomy and systematics
The woodlark was described by the Swedish naturalist Carl Linnaeus in 1758 in the tenth edition of his Systema Naturae and given the binomial name Alauda arborea. This binomial name is identical to the Latin name used in 1676 by English ornithologist Francis Willughby in his Ornithologiae libri tres. The woodlark is now placed in the genus Lullula that was introduced by the German naturalist Johann Jakob Kaup in 1829. The current genus name is onomatopoeic from French "Lulu", the name given to the bird by de Buffon. The French name, Alouette lulu, and the scientific name, Lullula arborea, are derived from the sound of its song.
Subspecies
There are two subspecies of woodlark:
- Northern woodlark (L. a. arborea) - <small>(Linnaeus, 1758)</small>: Found from northern, western and central Europe to western Russia and Ukraine
- Southern woodlark (L. a. pallida) - <small>Zarudny, 1902</small>: Found from southern Europe and north-western Africa through the Middle East to Iran and Turkmenistan, Crimea and Caucasus.
Description
thumb|right|alt=Drawing of a woodlark|Coloured lithograph by [[Magnus von Wright]]
This is a 13.5- to 15-cm-long bird, slightly smaller than the skylark being roughly 20% shorter. The woodlark is mainly brown above and pale below, but with a distinctive white superciliar meeting on the nape. It has a crest which is quite small and at most times inconspicuous. In flight it shows a short tail and short broad wings. The tail is tipped with white, but unlike the skylark, the tail sides and the rear edge of the wings are not edged with white.
A songbird, the woodlark has a melodious, warbling song often described onomatopoeically as a lu-lu-lu- In the same year, densities in East Germany ranged from 0.29 to 5.0 pairs per km<sup>2</sup> and between 0.1 and 0.25 pairs per km<sup>2</sup> in southern England, with more optimal habitats being more densely populated. However, populations fluctuated across Europe in the 1990s and 2000s and more up-to-date density figures are unavailable. The extent of the woodlark's range is England in the west, parts of northern Egypt to the south, Iran and Turkmenistan to the east and the Scandinavian Peninsula in the north. They prefer clearings in pine forests and heathland and like newly planted areas with pine saplings. The bird can also be found more rarely in urban areas. For example, in 1950 a pair were recorded on a main road near Putney Heath, London. Habitat loss is thought to be a major contributor to this, with dry grassland, fallow land, lowland heathland and pasture being lost to agriculture, abandonment and development across Northern Europe. Recent wildfires in England are also thought to have damaged the population, with some protected woodlark habitats having been destroyed.
In culture
The woodlark is commemorated in the works of two major poets. "The Woodlark", written by Gerard Manley Hopkins, departs from the standard tradition of British nature poetry by trying to transliterate the bird's song into made-up words.
The Scottish poet Robert Burns wrote of the bird's "melting art" in his poem "To the Woodlark". As there are currently no woodlarks in Scotland, and Burns never travelled south of Carlisle, many have speculated that Burns never came in contact with the bird and was in fact writing about the tree pipit, which was commonly referred to as the woodlark in Scotland. The woodlark's song is also thought to be melodious However, the woodlark has been spotted in Scotland on occasion and it is possible that Burns was writing about this bird. This is backed up by the entry of a minister from Clinic, Perthshire in the Old Statistical Account, which reads "The notes of the wood-lark are heard, delightful along the banks of the Lunan in spring and autumn; its nocturnal song has a dying cadence peculiarly melodious and has often been mistaken for the song of the Philomel [nightingale]."
Status
In Europe in 2004, the breeding population of woodlarks was estimated to number 1.3 million to 3.3 million breeding pairs. Europe accounts for 75-94% of the global population, meaning between 4.15 million and 13.2 million individuals in its world range. Populations of woodlark have fluctuated across Europe, with specific figures available for Britain showing these fluctuations. An estimated 400 breeding pairs were present in England in 1981. The woodlark has been categorised by the IUCN Red List of Threatened Species as being of least concern, meaning that it is not currently threatened with extinction.
