Risso's dolphin

Risso's dolphin (Grampus griseus) is the only species of the genus Grampus in the family Delphinidae (dolphins). Some of the most closely related species to these dolphins include: pilot whales (Globicephala spp.), pygmy killer whales (Feresa attenuata), melon-headed whales (Peponocephala electra), and false killer whales (Pseudorca crassidens). These dolphins grow to be about 10 ft in length and can be identified by heavy scarring that appears white.

Taxonomy

left|thumb|Skeleton of the [[holotype of Risso's dolphin, Paris National Museum of Natural History, 2025<sup>.</sup>]]

Risso's dolphin is named after Antoine Risso, whose study of the animal formed the basis of the recognized description by Georges Cuvier in 1812. The holotype referred to a specimen at the Muséum national d'histoire naturelle, an exhibit using preserved skin and skull obtained at Brest, France.

The type and sole species of the genus Grampus refers to Delphinus griseus Cuvier 1812. A proposition to name this genus Grampidelphis in 1933, when the taxonomic status of "blackfish" was uncertain, and conserving the extensive use of "Grampus" for the "killer" Orcinus orca", also suggested renaming this species (Grampidelphis exilis Iredale, T. & Troughton, E. le G. 1933). These were recognised as synonyms after publication of the Catalog of Whales (Hershkovitz, 1966).

thumb|A calf bears no scars, in contrast to its mother behind it.

Infants are dorsally grey to brown and ventrally cream-colored, with a white, anchor-shaped area between the pectorals and around the mouth. In older calves, the nonwhite areas darken to nearly black, and then lighten (except for the always dark dorsal fin). Linear scars mostly from social interaction eventually cover the bulk of the body; scarring is a common feature of male-to-male competition in toothed whales, but Risso's dolphin tend to be unusually heavily scarred. The pronounced appearance of these scars results from the lack of repigmentation, which may be advantageous, as a display that reduces further challenges from other males. Older individuals appear mostly white. Most individuals have two to seven pairs of teeth, all in the lower jaw. Like most dolphins, males are typically slightly larger than females. This species weighs , making it the largest species called "dolphin".

Range and habitat

left|thumb|A Risso's dolphin swims off [[Morro Bay.]]

thumb|A Risso's dolphin in the Gulf of Genoa

Risso's dolphins are found nearly worldwide, from cold and temperate to tropical waters, in the Indian, Pacific and Atlantic Oceans, as well as parts of the Baltic Sea, the Persian Gulf, and the Mediterranean, North, and Red Seas (excepting the Black Sea; however, a rare stranding was recorded in the Sea of Marmara in 2012).). Several sightings have been documented in Roskilde Fjord, in the waters of Lejre Vig, just off of the coast of Skjoldungernes Land National Park, Denmark. Analysis of Risso's dolphins found in the UK and the Mediterranean display variations in mitochondrial DNA. One possible reason for these differences could be the lack of interaction between individuals in the two locations.

In the Pacific, they range from French Polynesia west to Samoa, north to the Hawaiian Islands, as far as the Gulf of Alaska. However, they are absent from the waters of the western Pacific (off of Asia) beyond Futuna. They are quite common along the western coasts of British Columbia, the United States, and Mexico, continuing their range to the southern tip of Tierra Del Fuego. In the eastern Atlantic, they have been sighted as far south as the offshore waters of Liberia, Guinea, and Western Africa north through the Canary Islands and the Azores to southern Greenland. On the western Atlantic side, Risso's dolphins have been seen as far south as Guyana and Martinique; they can be found throughout much of the Caribbean Sea and the Gulf of Mexico to Florida and the Bahamas, and all along the American East Coast and the Canadian Maritime Provinces.

Since at least 2017, Risso's dolphins have begun to appear off of the subarctic Norwegian coast, as far north as Bleik's Canyon, off of Andøya. The repeated, regular sightings imply an expansion of their natural range. Possible explanations for this movement are a changing climate or varying water currents, as well as a northward migration of prey species or competition with other cetaceans, such as pilot whales.

Due to the low population density of the species, Risso's dolphins are widely considered difficult to establish an accurate estimate of population size in any given area.

A population is found off Santa Catalina Island, where they are sympatric with short-finned pilot whales (Globicephala macrorhynchus), and both species feed on the squid population. Although these species have not been seen to interact with each other, they take advantage of the commercial squid fishing that takes place at night. They have been seen by fishermen to feed around their boats. They also travel with other cetaceans. They surf the bow waves of gray whales, as well as ocean swells. These dolphins typically travel in groups of between 10 and 51, but can sometimes form "super-pods" reaching up to a few thousand individuals. Smaller, stable subgroups exist within larger groups. These groups tend to be similar in age or sex. Risso's experience fidelity towards their groups. Long-term bonds are seen to correlate with adult males. Younger individuals experience less fidelity and can leave and join groups. Mothers show a high fidelity towards a group of mother and calves,

Risso's dolphins are known to have a very active surface presence, often either displaying their tail flukes and pectoral fins, or slapping the surface of the water. They have also been known to engage in a behavior called spy-hopping, a common behavior in cetaceans where an individual vertically pokes its head out of the water. Recent studies have discussed the possibilities of spy-hopping as a sexual behavior, as it is typically only done in the presence of other individuals. Risso's dolphins do not require cutting teeth to process their cephalopod prey, which has allowed the species to evolve teeth as display weapons in mating conflicts.

Human interactions

Like other dolphins and marine animals, these dolphins getting caught in seine nets and gillnets have been documented across the globe.

In Ireland, though not apparently in England, Risso's dolphin was one of the royal fish, which by virtue of the royal prerogative, were the exclusive property of the English Crown.

A famed individual named Pelorus Jack was widely reported between 1888 and 1912, travelling with ships navigating the Cook Strait in New Zealand. A law protecting the animal was passed after a public outcry, renewed twice more, but suggested be invalid by its reference to fisheries acts that did not concern marine mammals.

Conservation

Risso's dolphin populations of the North, Baltic, and Mediterranean Seas are listed on Appendix II of the Convention on the Conservation of Migratory Species of Wild Animals (CMS), since they have an unfavourable conservation status or would benefit significantly from international co-operation organised by tailored agreements.

In addition, Risso's dolphin is covered by the Agreement on the Conservation of Small Cetaceans of the Baltic, North East Atlantic, Irish and North Seas (ASCOBANS), the Agreement on the Conservation of Cetaceans in the Black Sea, Mediterranean Sea and Contiguous Atlantic Area (ACCOBAMS), the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands Region (Pacific Cetaceans MoU) and the Memorandum of Understanding Concerning the Conservation of the Manatee and Small Cetaceans of Western Africa and Macaronesia (Western African Aquatic Mammals MoU).

Risso's dolphins are protected in the United States under the Marine Mammal Protection Act of 1992. Currently, Japan, Indonesia, the Solomon Islands, and The Lesser Antilles hunt Risso's dolphins. A juvenile male Risso's dolphin reportedly was stranded alive on the coast of Gran Canaria on 26 April 2019. This was the first documented case of capture myopathy and stress cardiomyopathy in a male juvenile Risso's dolphin that has received rehabilitation.

References

External links

Whale and Dolphin Conservation Society
ARKive Photographs, video
Voices in the Sea – Sound of the Risso's Dolphin