The ring ouzel (Turdus torquatus) is a mainly European member of the thrush family Turdidae. It is a medium-sized thrush, in length and weighing . The male is predominantly black with a conspicuous white crescent across its breast. Females are browner and duller than males, and young birds may lack the pale chest markings altogether. In all but the northernmost part of its range, this is a high-altitude species, with three subspecies breeding in mountains from Ireland east to Iran. It breeds in open mountain areas with some trees or shrubs, the latter often including juniper and other treeline conifers, rowan, bilberry, heather, and hairy alpenrose. It is a migratory bird, leaving the breeding areas to winter in southern Europe, North Africa and Turkey, typically in mountains with junipers. The typical clutch is 3–6 brown-flecked pale blue or greenish-blue eggs. They are incubated almost entirely by the female, with hatching normally occurring after 13 days. The altricial, downy chicks fledge in another 14 days and are dependent on their parents for about 12 days after fledging.

The ring ouzel is omnivorous, eating invertebrates, particularly insects and earthworms, some small vertebrates, and a wide range of fruit. Most animal prey is caught on the ground. During spring migration and the breeding season, invertebrates dominate the adult's diet and are also fed to the chicks. Later in the year, fruit becomes more important, particularly the common juniper.

With an extensive range and a large population, the ring ouzel is evaluated as least concern by the International Union for Conservation of Nature (IUCN). There are signs of decline in several countries; suspected causes including climate change, human disturbance, hunting and outdoor leisure activities. Loss of junipers may also be a factor in some areas. Natural hazards include predation by mammalian carnivores and birds of prey, and locally there may also be competition from other large thrushes such as the common blackbird, mistle thrush and fieldfare.

Etymology

"Ouzel" is an old name for the common blackbird, the word being cognate with the German . "Ouzel" may also be applied to a group of superficially similar but more distantly related birds, the dippers, the European representative of which is sometimes known as the water ouzel. "Ring Ouzel" was first used by John Ray in his 1674 Collection of English Words not Generally Used and became established with his 1678 book The Ornithology of Francis Willughby of Middleton in the County of Warwick. As with the English term, the scientific name also refers to the male's prominent white neck crescent, being derived from the Latin words , "thrush", and , "collared". Old and local names for the ring ouzel include "fell blackbird", "hill blackbird", "moor blackbird", "rock ouzel" and "mountain blackbird".

Taxonomy

The ring ouzel was first described by Carl Linnaeus under its current scientific name in his 1758 10th edition of Systema Naturae. He noted earlier descriptions by Francis Willughby and Eleazar Albin, both of whom gave it the name Merula torquata.

There are 104 species of medium to large thrushes in the genus Turdus. They are characterised by rounded heads, medium or longish pointed wings, and usually melodious songs.

A 2020 study of the genetics of Turdus

{| class="wikitable"

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! Image !! Scientific name !! Distribution !! Identification

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|200px<br />Cairngorm, Scotland || T.&nbsp;t.&nbsp;torquatus <br /><small>Linnaeus, 1758</small> || Breeds Ireland (rare), Great Britain to Scandinavia and the far northwest of Russia; winters from Spain south to northwest Africa and the Canary Islands || Black plumage with moderate pale fringes

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|200px<br />Hohe Tauern, Austria || T. t. alpestris <br /><small>(C.&nbsp;L.&nbsp;Brehm,&nbsp;1831)</small> || Breeds in the mountains of Iberia, the Pyrenees, the Alps, the Carpathians, and the Balkan Mountains, and with small outlying populations in the Atlas Mountains of Algeria, the Haut Vosges of Belgium, and the higher hills of northern Germany; winters in northern Africa, southern Europe and southern Turkey || All body feathering with strong pale fringes, giving a very 'scaly' appearance

|-

|200px<br />Artashavan, Armenia || T. t. amicorum <br /><small>E.&nbsp;Hartert,&nbsp;1923</small> || Breeds in central and eastern Turkey east through the Caucasus and Elburz Mountains to Turkmenistan; winters mainly in Iran and parts of Iraq || Blackest plumage with minimal pale fringing on the body feathers, but with very strong pale wing panel

|-

|}

Analysis of mitochondrial DNA samples from across Europe suggests that this species had a much broader distribution after the Last Glacial Period that ended about 11,700&nbsp;years ago than it does now.

Description

The ring ouzel is in length and weighs . The plumage of the male of the nominate race is entirely black except for a conspicuous white crescent on the breast, narrow greyish scaling on the upperparts and belly and pale edges to the wing feathers. The bill is yellow and the legs are greyish brown. The female resembles the male but is browner and with a duller breast band. Juveniles are like the female, but with a faint or non-existent breast crescent.

Males of T. t. alpestris have broader white scalloping (repeated small curves) on their underparts than T. t. torquatus, giving a distinctly scaly appearance below. The wing panel is also paler than in the nominate subspecies. Females are much as the nominate race, but with broad white fringes on the chin and throat. Adult ring ouzels undergo a complete moult after breeding from late June to early September, before their autumn migration. Juveniles have a partial moult between July and September, replacing their head, body and wing covert feathers.

The ring ouzel is extinct in Latvia and occurs only on migration in Denmark. It is a passage migrant in Syria and a vagrant to Iceland, Jordan, the Arabian Peninsula, Sudan, Kazakhstan, Mauritania, Svalbard and Jan Mayen. In Switzerland, ring ouzels breed on rugged upland slopes with heather, conifers, beech or hairy alpenrose at , The young are dependent on their parents for about 12 days after fledging. Around 36% of juveniles survive their first year, while the annual survival rate for adults is 47% for males and 37% for females. The main causes of death in northwest Europe are predation (9%), accidental human-related incidents (10%), and hunting, mainly in France (77%). and is key to the dispersal of the endemic Canary Islands juniper in the Canary Islands.

The young are mainly fed invertebrates, caterpillars and earthworms being major items where available. Although birds migrating in autumn use similar habitat to that used in spring, seasonal berries make up most of their diet, particularly elderberries, haws and, where available, juniper berries. common buzzard, common kestrel and Eurasian sparrowhawk, least weasel and stoat. Most deaths are of young juveniles, and birds hatched early in the season are more likely to survive than later broods. A Scottish study showed that raptors were responsible for 59% of deaths and mammals for 27%. In Romania, eggs were taken by red squirrels and spotted nutcrackers.

A study in the Carpathian Mountains found that a significant proportion of ring ouzels carried trombiculid mites. These mites commonly infect ground-feeding birds, and heavy infestations can cause birds to lose condition and stop feeding. The hard-bodied tick Ixodes festai commonly parasitises thrushes, including the ring ouzel. There is a record of this species carrying a Haemoproteus blood parasite.

Status and conservation

The ring ouzel has an extensive range, estimated at , and a large population, estimated at 600,000–2&nbsp;million individuals in Europe (which comprises 95% of the breeding range). The species is not believed to approach the thresholds for the population decline criteria of the IUCN Red List (i.e., declining more than 30% in ten years or three generations), and is therefore evaluated as least concern. The breeding population in Europe was estimated to be 299,000–598,000 pairs in 2019.

In the Alps, the density of breeding pairs can reach but is generally much lower with in Haute-Savoie, in the Jura Mountains, and in more open habitats in Britain.