Pinus sylvestris is a species of tree in the pine family Pinaceae that is native to Eurasia. It is commonly known as the Scots pine in English; it is also known as the Scotch pine in the United States, and occasionally called the Baltic pine or European red pine. It can readily be identified by its combination of fairly short, blue-green leaves and orange-red bark.

Names

Before the 18th century, the species was more often known as Scots fir or Scotch fir. Another, less common name is European redwood. or yellow deal. It is named after the deal, an obsolete unit of measurement for wood.

Description

thumb|upright|Young female cone

thumb|Pinus sylvestris forest in [[Sierra de Guadarrama, central Spain]]

Pinus sylvestris is a variety of ancient evergreen coniferous tree. A fossil seed cone of Pinus montana fossilis sent by the Naturmuseum Senckenberg to the Swedish Museum of Natural History was dated to the late Pliocene epoch (Reuverian stage ~ 2.6 million years ago). Its longevity might be due to its genetics. Genes of Scots Pine during the haploid stage of its life cycle are subject to stronger negative selection. Selective removal of alleles that contain potentially unhelpful mutations has the efficacy of efficient natural selection. and in trunk diameter when mature, exceptionally over tall and in trunk diameter on very productive sites. The tallest on record is a tree over 210 years old growing in Estonia which stands at . The lifespan is normally 150–300 years, with the oldest recorded specimens in Lapland, Northern Finland over 760 years.

The bark is thick, flaky and orange-red when young to scaly and gray-brown in maturity, sometimes retaining the former on the upper portion. They differ only minimally in morphology, but with more pronounced differences in genetic analysis and resin composition. Populations in westernmost Scotland are genetically distinct from those in the rest of Scotland and northern Europe, but not sufficiently to have been distinguished as separate botanical varieties. Trees in the far north of the range were formerly sometimes treated as var. lapponica, but the differences are clinal and it is not genetically distinct.

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! Image !! Varieties !!Description !! Distribution

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|120px|| Pinus sylvestris var. sylvestris ||Described above. || The bulk of the range, from Scotland and Spain to central Siberia.

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|120px|| Pinus sylvestris var. hamata ||Foliage more consistently glaucous all year, not becoming duller in winter; cones more frequently with a pyramidal apophysis. ||The Balkans, northern Turkey, Crimea, and the Caucasus.

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|120px|| Pinus sylvestris var. mongolica ||Foliage duller green, shoots gray-green; leaves occasionally up to 12 cm long. || Mongolia and adjoining parts of southern Siberia and northwestern China.

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|120px|| Pinus sylvestris var. nevadensis ||(not considered distinct from var. sylvestris by all authors) Kalenicz. Ex Kom. Cones often with thicker scales, but doubtfully distinguishable on morphology. ||The Sierra Nevada in southern Spain and possibly other Spanish populations

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|120px|| Pinus sylvestris var. cretacea || ||From border regions between Russia and Ukraine.

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Distribution and habitat

Pinus sylvestris is the only pine native to northern Europe, ranging from Western Europe to Eastern Siberia, south to the Caucasus Mountains and Anatolia, and north to well inside the Arctic Circle in Fennoscandia. In the north of its range, it occurs from sea level to , while in the south of its range it is a mountain tree, growing at altitude.

Britain and Ireland

thumb|Scattered survivors (two recently dead) of extensive deforestation at Glen Quoich, Scotland

The tree spread across Britain and Ireland after the Last Glacial Maximum. Pollen records show that pine was present locally in southern England by 9,000 years ago having entered from northeast France and that it had spread as far north as the Lake District and North Pennines 500 years later.

It was present in Ireland over 8,800 years ago but absent from Wales at that time which suggests that the pine in Ireland had a separate Iberian origin or contained surviving populations, although evidence towards its survival is lacking.

In Britain it now occurs naturally only in Scotland. Historical and archaeological records indicate that it also occurred in Wales and England until about 300–400 years ago, becoming extinct there due to over-exploitation and grazing; it has been re-introduced in these countries. Similar historical extinction and re-introduction applies to Ireland, Denmark and the Netherlands. Whether it truly became extinct in England is unknown. It has been speculated that it may have survived wild long enough for trees used in cultivation in England to derive from native (rather than imported) sources. Shakespeare (in Richard II) was familiar with the species in the 1590s, as was Evelyn in the early 1660s (Sylva), both around the time when the pine was thought to become extinct in England, but when landowners were also beginning ornamental and forestry planting.of this ancient forest remain, the main surviving remnants being at Abernethy Forest, Glen Affric, Rothiemurchus Forest, and the Black Wood of Rannoch. Plans are currently in progress to restore at least some areas and work has started at key sites.

Uses

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thumb|Scots pine forest in [[Estonia]]

Pinus sylvestris is an important tree in forestry. The wood is used for pulp and sawn timber products. A seedling stand can be created by planting, sowing, or natural regeneration. Commercial plantation rotations vary between 50 and 120 years, with longer rotations in northeastern areas where growth is slower.

In Scandinavian countries, the pine was used for making tar in the preindustrial age. Some active tar producers still exist, but that industry has almost ceased. which has a hemp-like appearance, but with a tighter, softer texture.

The pine has also been widely planted in New Zealand and much of the colder regions of North America; it was one of the first trees introduced to North America, in about 1600. It is listed as an invasive species in some areas there, including Ontario, Michigan. It has been widely used in the United States for the Christmas tree trade, and was one of the most popular Christmas trees from the 1950s through the 1980s. It remains popular for that usage, though it has been eclipsed in popularity, by such species as Fraser fir, Douglas-fir, and others. Despite its invasiveness in parts of eastern North America, the pine does not often grow well there, partly due to climate and soil differences between its native habitat and that of North America, and partly due to damage by pests and diseases; the tree often grows in a twisted, haphazard manner if not tended to (as they are in the Christmas tree trade).

Previously, the pine was grown in and used extensively by the coal mining regions of Flanders, Belgium. It was used to fortify tunnels, primarily because it would make a cracking sound when in need of replacement. Large patches of forest, mostly containing the species, are still scattered over the countryside.

Cultivars

Several cultivars are grown for ornamental purposes in parks and large gardens, of which 'Aurea', 'Beuvronensis', 'Frensham', and 'Gold Coin' have gained the Royal Horticultural Society's Award of Garden Merit.

In culture

The Scots pine is the plant badge of Clan Gregor and Clan Farquharson. It is the national tree of Scotland.

<gallery mode="packed">

Pinus sylvestris cones pl.jpg|Leaves and cones, Poland

Pinus syvestris seedling.jpg|Seedling with flattish, unfascicled leaves

Pinus sylvestris wood ray section 1 beentree.jpg|Wood

Pine log cross-sections, Teutendorf, 2016-09-17.jpg|Cross-sections of pine logs from a thinning

Pinus Sylvestris 8407.jpg|Bark on a mature specimen

Strom roka borovica velke borove 03.jpg|Looking up in the branch structure

Pine releasing pollen into the wind in Tuntorp 1.jpg|Pine releasing pollen on a windy day in Sweden

Seed pinus sylvestris 1 beentree B.jpg|Seeds

</gallery>

See also

  • Pinus × rhaetica

References

  • Pinus sylvestris - distribution map, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN)