Cingulata, part of the superorder Xenarthra, is an order of armored New World placental mammals. The armadillos, whose species are split between the families Dasypodidae and Chlamyphoridae, are the only surviving members of the order. Two groups of cingulates much larger than extant armadillos (maximum body mass of 45 kg (100 lb) in the case of the giant armadillo) existed until recently: pampatheriids, which reached weights of up to 200 kg (440 lb) and chlamyphorid glyptodonts, which attained masses of 2,000 kg (4,400 lb) or more.
The cingulate order originated in South America during the Paleocene epoch about 66 to 56 million years ago, and due to the continent's former isolation remained confined to it during most of the Cenozoic. However, the formation of a land bridge allowed members of all three families to migrate to southern North America during the Pliocene or early Pleistocene as part of the Great American Interchange. After surviving for tens of millions of years, all of the pampatheriids and giant glyptodonts apparently died out during the Quaternary extinction event at the beginning of the Holocene, along with much of the rest of the regional megafauna, shortly after the colonization of the Americas by Paleo-Indians.
Description
Armadillos have dorsal armor that is formed by osteoderms, plates of dermal bone covered in relatively small, overlapping keratinized epidermal scales called "scutes". Most species have rigid shields over the shoulders and hips, with three to nine bands separated by flexible skin covering the back and flanks.
Pampatheres also had shells that were flexible due to three movable lateral bands of osteoderms. Euphractinae is unique for speciations towards carnivory, culminating in the macropredatory genus Macroeuphractus.
Classification
thumb|right|[[Holmesina septentrionalis (Barcelona)]]
thumb|right|[[Nine-banded armadillo, D. novemcinctus (Smithsonian)]]
thumb|right|[[Glyptodon clavipes (Berlin)]]
thumb|right
The taxonomic table below follows the results of a phylogenetic analysis published by Delsuc et al., 2016. While glyptodonts have traditionally been considered stem-group cingulates outside the group that contains modern armadillos, this 2016 study conducted an analysis of Doedicurus mtDNA and found that it was, in fact, nested within the modern armadillos as the sister group of a clade consisting of Chlamyphorinae and Tolypeutinae.
Order Cingulata
- Family †Peltephilidae <small>Ameghino 1894</small>
- Genus †Anantiosodon <small>Ameghino 1891</small>
- Genus †Epipeltephilus <small>Ameghino 1904</small>
- Genus †Parapeltecoelus <small>Bordas 1938</small>
- Genus †Peltecoelus <small>Ameghino 1902</small>
- Genus †Peltephilus <small>Ameghino 1887</small> (Horned armadillo)
- Genus †Ronwolffia <small>Shockey 2017</small>
- Family †Paleopeltidae <small>Ameghino 1895</small>
- Genus †Palaeopeltis <small>Ameghino 1895</small>
- Family †Pampatheriidae
- Genus ?†Machlydotherium <small>Ameghino 1902</small>
- Genus †Holmesina <small>Simpson 1930</small>
- Genus †Kraglievichia <small>Castellanos 1927</small>
- Genus †Machlydotherium
- Genus †Pampatherium <small>Ameghino 1875 ex Gervais & Ameghino 1880</small>
- Genus †Scirrotherium <small>Edmund & Theodor 1997</small>
- Genus †Tonnicinctus <small>Góis et al. 2015</small>
- Genus †Vassallia <small>Castellanos 1927</small> [Plaina <small>Castellanos 1937</small>]
- Genus †Yuruatherium <small>Ciancio et al. 2012</small>
- Family †Pachyarmatheriidae <small>Fernicola et al. 2018</small>
- Genus †Neoglyptatelus <small>Carlini, Vizcaíno & Scillato-Yané 1997</small>
- Genus †Pachyarmatherium <small>Downing & White 1995</small>
- Family Dasypodidae (long-nosed armadillos)
- Genus †Acantharodeia
- Genus †Amblytatus
- Genus †Archaeutatus
- Genus †Astegotherium
- Genus †Barrancatatus
- Genus †Chasicotatus
- Genus †Chorobates
- Genus †Coelutaetus
- Genus †Eocoleophorus
- Genus †Epipeltecoelus
- Genus †Eutatus
- Genus †Hemiutaetus
- Genus †Isutaetus
- Genus †Lumbreratherium
- Genus †Macrochorobates
- Genus †Mazzoniphractus
- Genus †Meteutatus
- Genus †Pedrolypeutes
- Genus †Prodasypus
- Genus †Proeutatus
- Genus †Prostegotherium
- Genus †Pucatherium
- Genus †Punatherium
- Genus †Stegotherium
- Genus †Stenotatus
- Genus †Utaetus
- Genus †Vetelia
- Subfamily Dasypodinae
- Genus †Anadasypus
- Genus Dasypus
- Genus †Nanoastegotherium
- Genus †Parastegosimpsonia
- Genus †Pliodasypus
- Genus †Propraopus
- Genus †Riostegotherium
- Genus †Stegosimpsonia
- Family Chlamyphoridae: glyptodonts and other armadillos
- Subfamily Chlamyphorinae: fairy armadillos
- Genus Calyptophractus
- Genus Chlamyphorus
- Subfamily Euphractinae: hairy, six-banded and pichi armadillos
- Genus Chaetophractus
- Genus †Doellotatus
- Genus Euphractus
- Genus †Macroeuphractus
- Genus †Proeuphractus
- Genus †Paleuphractus
- Genus Zaedyus
- Subfamily †Glyptodontinae: glyptodonts
- Genus †Doedicurus
- Genus †Glyptodon
- Genus †Glyptotherium
- Genus †Hoplophorus
- Genus †Panochthus
- Genus †Parapropalaehoplophorus
- Genus †Plaxhaplous
- Subfamily Tolypeutinae: giant, three-banded and naked-tailed armadillos
- Genus Cabassous
- Genus †Kuntinaru
- Genus Priodontes
- Genus Tolypeutes
